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Ownership Document
|
| Schema Version: X0508 |
| Document Type: 4 |
| Period of Report: 11/17/23 |
| Issuer: |
| Issuer CIK: 1497253 |
| Issuer Name: ORGANOVO HOLDINGS, INC. |
| Issuer Trading Symbol: ONVO |
| Reporting Owner: |
| Reporting Owner ID: |
| | Owner CIK: 1536583 |
| | Owner Name: Murphy Keith |
| Reporting Owner Address: |
| | Owner Street 1: 11555 SORRENTO VALLEY ROAD, SUITE 100 |
| | Owner Street 2: |
| | Owner City: SAN DIEGO |
| | Owner State: CA |
| | Owner ZIP Code: 92121 |
| | Owner State Description: |
| Reporting Owner Relationship: |
| | Is Director? Yes |
| | Is Officer? Yes |
| | Is Ten Percent Owner? No |
| | Is Other? No |
| | Officer Title: Executive Chairman |
| Aff 10b5 One? No |
| Non-Derivative Table: |
| Non-Derivative Transaction: |
| | Security Title: |
| | | Value: Common Stock |
| | Transaction Date: |
| | | Value: 11/17/23 |
| | Transaction Coding: |
| | | Transaction Form Type: 4 |
| | | Transaction Code: A |
| | | Equity Swap Involved? No |
| | Transaction Amounts: |
| | | Transaction Shares: |
| Value: 19,607 |
| Footnote ID: F1 |
| | | Transaction Price Per Share: |
| Value: 0.00 |
| | | Transaction Acquired-Disposed Code: |
| Value: A |
| | Post-Transaction Amounts: |
| | | Shares Owned Following Transaction: |
| Value: 113,034 |
| | Ownership Nature: |
| | | Direct or Indirect Ownership: |
| Value: D |
| Footnotes: |
| Footnote - F1: The reported securities are represented by restricted stock units, which shall vest on the earlier of November 17, 2024 or the date of the next annual meeting of stockholders held by the Issuer, subject to acceleration in the event of a change of control. |
| Owner Signature: |
| Signature Name: /s/ Keith Murphy |
| Signature Date: 11/17/23 |
