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Maroun Richard E – ‘4’ for 9/5/19 re: Organovo Holdings, Inc.

On:  Monday, 9/9/19, at 7:00pm ET   ·   For:  9/5/19   ·   Accession #:  1209191-19-48962   ·   File #:  1-35996

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  As Of               Filer                 Filing    For·On·As Docs:Size             Issuer                      Filing Agent

 9/09/19  Maroun Richard E                  4                      2:8K   Organovo Holdings, Inc.           DONNELLEY FIN… File16/FA

Statement of Changes in Beneficial Ownership of Securities by an Insider   —   Form 4   —   SEA’34
Filing Table of Contents

Document/Exhibit                   Description                      Pages   Size 

 1: 4           Form 4 Submission -- doc4.xml/3.6                   HTML      3K 
 2: EX-24.4_871788  POA Document                                       1      6K

‘4’   —   Form 4 Submission — doc4.xml/3.6



        

This ‘4’ Document is an XML Data File that may be rendered in various formats:

  Form 4    –   Plain Text   –  SEC Website  –  EDGAR System  –    XML Data    –  <?xml?> File
 
 
XML Data rendering:  Form 4 Submission
 
                                                                                                                                                                                
<?xml version="1.0" encoding="windows-1252"?>
<ownershipDocument>
<schemaVersion> X0306 </schemaVersion>
<documentType> 4 </documentType>
<periodOfReport> 2019-09-05 </periodOfReport>
<notSubjectToSection16> 0 </notSubjectToSection16>
<issuer>
<issuerCik> 0001497253 </issuerCik>
<issuerName> ORGANOVO HOLDINGS, INC. </issuerName>
<issuerTradingSymbol> ONVO </issuerTradingSymbol>
</issuer>
<reportingOwner>
<reportingOwnerId>
<rptOwnerCik> 0001364590 </rptOwnerCik>
<rptOwnerName> Maroun Richard E </rptOwnerName>
</reportingOwnerId>
<reportingOwnerAddress>
<rptOwnerStreet1> 6275 NANCY RIDGE DRIVE, SUITE 110 </rptOwnerStreet1>
<rptOwnerStreet2></rptOwnerStreet2>
<rptOwnerCity> SAN DIEGO </rptOwnerCity>
<rptOwnerState> CA </rptOwnerState>
<rptOwnerZipCode> 92121 </rptOwnerZipCode>
<rptOwnerStateDescription></rptOwnerStateDescription>
</reportingOwnerAddress>
<reportingOwnerRelationship>
<isDirector> 1 </isDirector>
<isOfficer> 0 </isOfficer>
<isTenPercentOwner> 0 </isTenPercentOwner>
<isOther> 0 </isOther>
</reportingOwnerRelationship>
</reportingOwner>
<derivativeTable>
<derivativeTransaction>
<securityTitle>
<value> Stock Option (Right to Buy) </value>
</securityTitle>
<conversionOrExercisePrice>
<value> 0.266 </value>
</conversionOrExercisePrice>
<transactionDate>
<value> 2019-09-05 </value>
</transactionDate>
<deemedExecutionDate></deemedExecutionDate>
<transactionCoding>
<transactionFormType> 4 </transactionFormType>
<transactionCode> A </transactionCode>
<equitySwapInvolved> 0 </equitySwapInvolved>
</transactionCoding>
<transactionTimeliness>
<value></value>
</transactionTimeliness>
<transactionAmounts>
<transactionShares>
<value> 52000 </value>
</transactionShares>
<transactionPricePerShare>
<value> 0.00 </value>
</transactionPricePerShare>
<transactionAcquiredDisposedCode>
<value> A </value>
</transactionAcquiredDisposedCode>
</transactionAmounts>
<exerciseDate>
<footnoteId id="F1"/>
</exerciseDate>
<expirationDate>
<value> 2029-09-05 </value>
</expirationDate>
<underlyingSecurity>
<underlyingSecurityTitle>
<value> Common Stock </value>
</underlyingSecurityTitle>
<underlyingSecurityShares>
<value> 52000 </value>
</underlyingSecurityShares>
</underlyingSecurity>
<postTransactionAmounts>
<sharesOwnedFollowingTransaction>
<value> 52000 </value>
</sharesOwnedFollowingTransaction>
</postTransactionAmounts>
<ownershipNature>
<directOrIndirectOwnership>
<value> D </value>
</directOrIndirectOwnership>
</ownershipNature>
</derivativeTransaction>
</derivativeTable>
<footnotes>
<footnote id="F1"> The option represents an automatic annual award granted pursuant to the Issuer's non-employee director compensation plan. The option shares vest and become exercisable in full on the earlier of (i) September 5, 2020 or (ii) immediately prior to commencement of the Issuer's next annual stockholder meeting. </footnote>
</footnotes>
<remarks></remarks>
<ownerSignature>
<signatureName> /s/ Jennifer Bush, as attorney-in-fact </signatureName>
<signatureDate> 2019-09-09 </signatureDate>
</ownerSignature>
</ownershipDocument>


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Filing Submission 0001209191-19-048962   –   Alternative Formats (Word / Rich Text, HTML, Plain Text, et al.)
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