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<?xml version="1.0" encoding="windows-1252"?> |
<ownershipDocument> |
| <schemaVersion> X0306 </schemaVersion> |
| <documentType> 4 </documentType> |
| <periodOfReport> 2013-10-07 </periodOfReport> |
| <notSubjectToSection16> 0 </notSubjectToSection16> |
| <issuer> |
| | <issuerCik> 0001497253 </issuerCik> |
| | <issuerName> ORGANOVO HOLDINGS, INC. </issuerName> |
| | <issuerTradingSymbol> ONVO </issuerTradingSymbol> |
| | </issuer> |
| <reportingOwner> |
| | <reportingOwnerId> |
| | | <rptOwnerCik> 0001549671 </rptOwnerCik> |
| | | <rptOwnerName> David Eric </rptOwnerName> |
| | | </reportingOwnerId> |
| | <reportingOwnerAddress> |
| | | <rptOwnerStreet1> 6275 NANCY RIDGE DRIVE, SUITE 110 </rptOwnerStreet1> |
| | | <rptOwnerStreet2></rptOwnerStreet2> |
| | | <rptOwnerCity> SAN DIEGO </rptOwnerCity> |
| | | <rptOwnerState> CA </rptOwnerState> |
| | | <rptOwnerZipCode> 92121 </rptOwnerZipCode> |
| | | <rptOwnerStateDescription></rptOwnerStateDescription> |
| | | </reportingOwnerAddress> |
| | <reportingOwnerRelationship> |
| | | <isDirector> 0 </isDirector> |
| | | <isOfficer> 1 </isOfficer> |
| | | <isTenPercentOwner> 0 </isTenPercentOwner> |
| | | <isOther> 0 </isOther> |
| | | <officerTitle> Chief Strategy Officer </officerTitle> |
| | | </reportingOwnerRelationship> |
| | </reportingOwner> |
| <nonDerivativeTable> |
| | <nonDerivativeTransaction> |
| | | <securityTitle> |
| <value> Common Stock </value> |
| </securityTitle> |
| | | <transactionDate> |
| <value> 2013-10-07 </value> |
| </transactionDate> |
| | | <transactionCoding> |
| <transactionFormType> 4 </transactionFormType> |
| <transactionCode> S </transactionCode> |
| <equitySwapInvolved> 0 </equitySwapInvolved> |
| <footnoteId id="F1"/> |
| </transactionCoding> |
| | | <transactionAmounts> |
| <transactionShares> |
| <value> 63000 </value> |
| </transactionShares> |
| <transactionPricePerShare> |
| <value> 5.8564 </value> |
| <footnoteId id="F2"/> |
| </transactionPricePerShare> |
| <transactionAcquiredDisposedCode> |
| <value> D </value> |
| </transactionAcquiredDisposedCode> |
| </transactionAmounts> |
| | | <postTransactionAmounts> |
| <sharesOwnedFollowingTransaction> |
| <value> 731306 </value> |
| <footnoteId id="F3"/> |
| </sharesOwnedFollowingTransaction> |
| </postTransactionAmounts> |
| | | <ownershipNature> |
| <directOrIndirectOwnership> |
| <value> D </value> |
| </directOrIndirectOwnership> |
| </ownershipNature> |
| | | </nonDerivativeTransaction> |
| | </nonDerivativeTable> |
| <footnotes> |
| | <footnote id="F1"> The sale of shares was effected pursuant to a Rule 10b5-1 trading plan adopted by the Reporting Person. </footnote> |
| | <footnote id="F2"> This transaction was executed in multiple sales through a sale order executed by a broker-dealer at prices ranging from $5.74 to $6.08. The price reported in this column reflects the weighted average sale price. The Reporting Person will provide upon request to the SEC staff, the Issuer, or a security holder of the Issuer, full information regarding the number of shares sold at each separate price. </footnote> |
| | <footnote id="F3"> This number does not include 700,000 shares of common stock underlying outstanding options and 20,000 shares of common stock underlying warrants held by Mr. David; post-transaction Mr. David continues to beneficially own or have a right to acquire 1,451,306 shares of common stock. </footnote> |
| | </footnotes> |
| <ownerSignature> |
| | <signatureName> /s/ Eric David </signatureName> |
| | <signatureDate> 2013-10-09 </signatureDate> |
| | </ownerSignature> |
| </ownershipDocument> |